☀️ Photosynthesis in Higher Plants

1. Introduction and Site

Physicochemical process by which green plants use light energy to synthesize organic compounds. Occurs in Chloroplasts. (Light reactions in grana, Dark reactions in stroma).

Main pigment: Chlorophyll a. Accessory pigments: Chlorophyll b, Xanthophylls, Carotenoids (protect from photo-oxidation).

2. Light Reaction (Photochemical Phase)

Involves light absorption, water splitting, oxygen release, and formation of ATP and NADPH.

Photosystems

PS II (P680): Involved in Non-cyclic photophosphorylation. Splitting of water occurs here.

PS I (P700): Involved in both Cyclic and Non-cyclic photophosphorylation.

Chemiosmotic Hypothesis: ATP synthesis is linked to the development of a proton gradient across the thylakoid membrane.

3. Dark Reaction (Biosynthetic Phase)

Does not require light directly, but depends on the products of light reaction (ATP, NADPH) to fix CO₂.

C3 and C4 Pathways

C3 Pathway (Calvin Cycle): Primary CO₂ acceptor is RuBP (5C). First stable product is 3-PGA (3C). Key enzyme: RuBisCO.

C4 Pathway (Hatch-Slack): Occurs in plants adapted to dry tropical regions (Maize, Sugarcane). Primary CO₂ acceptor is PEP (3C). First product is OAA (4C). Show Kranz anatomy. Prevents photorespiration.

4. Photorespiration

A wasteful process in C3 plants where RuBisCO acts as an oxygenase (instead of carboxylase) binding to O₂. It releases CO₂ and consumes ATP. No sugar is formed.